Evolution

Like Noth, the crocodile had woken early, disturbed from its slumber by the brightness of the day. As it rose from the lake, bits of ice tumbled from its back. With a single graceful motion it clamped its jaws on the frozen meso: frost crackled, bones crunched. Then the croc slid backward into the water, dragging the carcass effortlessly, making barely a sound.

The crocodile was hungry.

Before the comet the largest animals in each of the world’s ecologies had been reptilian: the plesiosaurs and ichthyosaurs in the oceans, the dinosaurs on land, and the crocodilians in fresh water. The disaster had wiped away these great families, and in their empty realms they would soon be replaced by functionally equivalent mammals — all save the crocodiles.

The freshwater environment had always been a difficult place to live. While the supply of plant material on land and in the sea was pretty reliable in space and time, freshwater environments were very variable. Erosion, abrasion, silting, flood, drought, and extremes of water quality were all hazards.

But the crocodiles — and other enduring freshwater species like turtles — were resilient. Some learned to walk overland in search of water. Others could take to the sea. Or they would just bury themselves eight or ten meters deep in the mud, and wait for the next cloudburst. And as for food, even during the worst of the killings on land and in the sea, they would subsist off the nutrients that continued to leach off the corpse-littered land, a "brown" food chain that persisted long after the green, growing things and the creatures that browsed them had died.

In this way the crocodiles had survived across a hundred and fifty million years, through extraterrestrial impacts, glaciation pulses, sea level changes, tectonic upheavals, and competition from successive dynasties of animals.

After all this time they were still capable of evolutionary novelty. Briefly, after the comet impact, the top predators around the water courses had been crocodile cousins with long legs and hooflike claws. They had been a nightmare, running predatory crocodiles capable of chasing down animals as large as small horses. Crocodiles had even adapted to survive here at the pole, where the sun didn’t shine for months on end; they would simply wait out the winter months in deep hibernation.

Unlike the dinosaurs, unlike the plesiosaurs, the crocodiles would not be forced out of their freshwater niches by upstart mammals: not now, not ever.

Noth had lost the meso carcass, but some scraps of flesh and crushed maggots were smeared over the ground where it had lain. Hungrily he licked at the frozen ground.

At last the days of breeding arrived.

The females of the troop gathered in the branches of one tall conifer. They were feeding on ripe young fruit, cramming their bodies with the resources they would need to survive the drain of motherhood to come. The females were loosely marshaled by the more senior among them, including Big and Biggest. Right was among them. She had survived her first winter. She was filling out quickly, and when her scrappy winter fur had blown away she had emerged as a small but elegantly built adult, ready to mate.

The Emperor himself was among his female subjects. He moved from one to the other, heroically humping. Already he had been accepted by Biggest twice, and had deflowered an unprotesting Right. Now he was taking Big. She was bent over, clinging to a low branch, her head tucked between her knees, her tail uplifted. The Emperor was behind her, arms wrapped around her waist, hips thrusting with a rapidity born of exhaustion and urgency.

This was the day toward which the Emperor had worked all year, and now was the time for him to spend all of his authority and energy by covering as many of the females as possible.

But the Emperor was already tiring. And this female troop was only one of several in the wider territory he commanded.

In this ferociously seasonal place, baby rearing had to be squeezed into a drastically short period, so that offspring were produced when food was abundant and their new mothers could eat enough to produce plenty of milk. Any female who mated outside the breeding season was unlikely to see her offspring survive to adulthood. And any male who missed the chance to mate with a fertile female would have to endure a whole year of hardship, danger, and privation before getting another chance.

For the notharctus, the breeding season was just forty-eight hours long. It was a frantic time.