Stern and Susman did in fact claim that a few of the Laetoli footprints gave signs of toes longer than in humans. Tuttle (1985, p. 132) admitted that “the right foot of G-1 sometimes left peculiar marks distal to the toe tips.” To Stern and Susman, the marks forward of the “toe tips” represented the actual toe tips of uncurled toes. But Tuttle had another explanation for the marks. He wrote: “These are best explained by . . . the tendency for G-1 to drag its foot on lift off probably due to pathology of the lower limb” (Tuttle 1985, p. 132). The fact that the peculiar markings appeared only on one foot of one individual, and then only sometimes, lends support to Tuttle’s explanation.
Stern and Susman (1983) also suggested that the Laetoli prints did not have a deep rounded impression at the base of the big toe, representing the ball of the foot in humans. They regarded this as evidence that the foot that made the prints was not human. But Tuttle (1985, p. 132) observed that “humans commonly leave prints devoid of these features as may be seen in prints on the beach.” And, as we have seen, Robbins (1987, p. 501) said the prints she studied did have a “humanlike” ball region.
Directly challenging Johanson, White, Latimer, and Lovejoy, who asserted
Tim White, for example, published a study (White and Suwa 1987) of the Laetoli prints in which he disputed Tuttle’s contention that their maker was a hominid more advanced than
White asserted: “there is not a single shred of evidence among the 26 hominid individuals in the collection of over 5,000 vertebrate remains from Laetoli that would suggest the presence of a more advanced Pliocene hominid at this site” ( White and Suwa 1987, p. 496). But, as we have seen in our review of African hominid fossils, there are in fact a few “shreds” of evidence for the presence of
Like Tuttle, White rejected the curled-toe hypothesis of Stern and Susman. Instead, White tried to fit the foot of
According to White, the best tracks at Laetoli were in the G-1 trail, representing the smallest of the three individuals of the G group. Even White admitted that the phalanges of AL 333-115 were “obviously incompatible with the G-1 tracks” (White and Suwa 1987, p. 497). Stern and Susman, and Tuttle, found them incompatible with any of the tracks. White, however, pointed out that the AL 333-115 individual represented one of the larger, presumably male, members of the First Family group and proposed that the foot of Lucy, one of the smaller, female individuals, might have fitted the G-1 Laetoli prints.
But the only bones recovered from Lucy’s foot were an ankle bone and two toe bones. White therefore decided to use a partial
White predicted that “the discovery of a complete foot skeleton at Hadar or Laetoli will conform in its basic proportions with the reconstruction described in this paper” ( White and Suwa 1987, p. 512). But this prediction remains to be fulfilled. It is interesting that the most complete